diff --git a/sqlparse/keywords.py b/sqlparse/keywords.py
index dd6e5d15..c5c65688 100644
--- a/sqlparse/keywords.py
+++ b/sqlparse/keywords.py
@@ -132,6 +132,7 @@ def find_delimited_spans(text):
     (r"`(``|[^`])*`", tokens.Name),
     (r"´(´´|[^´])*´", tokens.Name),
 
+    (r'@\?', tokens.Operator),
     (r'\?', tokens.Name.Placeholder),
     (r'%(\(\w+\))?s', tokens.Name.Placeholder),
     (r'(?<!\w)[$:?]\w+', tokens.Name.Placeholder),
diff --git a/tests/test_format.py b/tests/test_format.py
index 93495067..5253911c 100644
--- a/tests/test_format.py
+++ b/tests/test_format.py
@@ -363,6 +363,15 @@ def test_bools(self):
         assert self.formatter(
             sql) == 'select * from table where a && b or c || d'
 
+    def test_json_path_operator(self):
+        sql = 'select data@?? from table'
+        assert self.formatter(sql) == 'select data @? ? from table'
+
+    @pytest.mark.parametrize('operator', ['||', '|', '&'])
+    def test_operator_after_placeholder(self, operator):
+        sql = f'select ?{operator}? from table'
+        assert self.formatter(sql) == f'select ? {operator} ? from table'
+
     def test_nested(self):
         sql = 'select *, case when a-b then c end from table'
         assert self.formatter(
diff --git a/tests/test_parse.py b/tests/test_parse.py
index 67168410..d1c41624 100644
--- a/tests/test_parse.py
+++ b/tests/test_parse.py
@@ -632,8 +632,8 @@ def test_configurable_regex():
 @pytest.mark.parametrize('sql', [
     '->', '->>', '#>', '#>>',
     '@>', '<@',
-    # leaving ? out for now, they're somehow ambiguous as placeholders
-    # '?', '?|', '?&',
+    # Question-mark prefixes remain ambiguous with placeholders.
+    '@?', '@@',
     '||', '-', '#-'
 ])
 def test_json_operators(sql):